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PUBLICATIONS

IN THE FIELD, 1990-2024

2024 Rafiq, K., Jordan, N.R., McNutt, J.W., Neelo, J., Attias, N., Boersma , P.D., Palmer, M., Ruesink, J., and Abrahms, B. . Removing fieldwork barriers is critical for advancing ecology. Trends in Ecology and Evolution. https://doi.org/10.1016/j.tree.2024.10.003

2024 Bennitt, E., Bartlam-Brooks, H., Hubel, T., Jordan, N., McNutt, J., Wilson, A. Proactive cursorial and ambush predation risk avoidance in four African herbivore species. Ecology and Evolution. (in press).

2024 Claase, M., Cherry, M.I., Hofmann, D.D., Apps, P. J., McNutt, J.W., & Jordan, N. R . Patterns of scent-marking by African wild dogs, Lycaon pictus, at shared marking sites. Animal Behaviour 207:77-86.

2024 Rafiq, K., Jordan, N.R., McNutt, J.W., Neelo, J., Attias, N., Boersma , P.D., Palmer, M., Ruesink, J., and Abrahms, B.  The value of field research in academia. Science, 384:6698 (855-856).

2024 Hansen, K.W., Ranc, N.; Morgan, J.; Jordan, N.; McNutt, J. W.; Wilson, A.; Wilmers, C. How territoriality and sociality influence the habitat selection and movement of a large carnivore. Ecology and Evolution 14(4): e11217. 

2024 Hofmann, D., Behr, D., McNutt, J.; Ozgul, A.; Cozzi, G. Dispersal and Connectivity in Increasingly Extreme Climatic Conditions. Global Change Biology 30(5): e17299.

2024  West, L., Rafiq, K., Converse, S. J., Wilson, A. M.  Jordan, N.R., Golabek,K.A. , McNutt, J. W.,  Abrahms, B. Droughts reshape apex predator space use and intraguild overlap. Journal of Animal Ecology. http://doi.org/10.1111/1365-2656.14192

2024 Hansen, KW, Jordan, NR, Claase, MJ, Suraci, JP, McNutt, JW, Dhruv, A, and Wilmers, CC. Experimental Modification of African Wild Dog Movement and Behavior Using Translocated Conspecific Scent. Biological Conservation 294: 110645.

2024 Claase, M.J., Cherry, M.I., McNutt, J.W., Apps, P.J., & Jordan, N.R. Size matters: natural experiments suggest the dear enemy effect is moderated by pack size in African wild dogs. Ecology and Evolution 14(4): e11298.

2023 Behr, D., Hofmann, D., McNutt, J.; Ozgul, A.; Cozzi, G. Higher mortality is not a universal cost of dispersal: Case of the African wild dog. The American Naturalist 202:5, 616-629

2023 Hofmann, D.; Behr, D.; McNutt, J.; Ozgul, A.; Cozzi, G. A Three-Step Approach for Assessing Landscape Connectivity via Simulated Dispersal: African Wild Dog Case Study. Landscape Ecology 38(4): 981–98.

2023 Apps, P., & McNutt, J.W.  Reactions of Free-Ranging African Carnivores to Single Components of Carnivore Odours: Evidence that Mammalian Semiochemicals Are Not All Irreducibly Complex. In: Schaal, B., Rekow, D., Keller, M., Damon, F. (eds) Chemical Signals in Vertebrates 15. CSiV 2021. Springer, Cham.

2023 Cozzi, G., Reilly, M., Abegg, D., Behr, D.M., Brack, P., Claase, M.J., Holmberg,J., Hofmann,D.D., Kalil, P., Ndlovu, S., Neelo, J., & McNutt, J.W. . An AI-based platform to investigate African large carnivore dispersal and demography across 2 broad landscapes:  A case study and future directions using African wild dogs. Afr. J. Ecol., 00, 1–9.


2023 Woodroffe, R., Abrahms, B., English,H., Jumbame, K., Lindena, J., Ngatiaa, D., Rabaiotti, D., & McNutt, J.W. African wild dogs are hot and hungry: Response to Creel et al.. Biological Conservation, 284. 110198.

2023 Rafiq, K., Jordan, N.R., Golabek, K, McNutt, J.W, Wilson, A, & Abrahms, B. Increasing ambient temperatures trigger shifts in activity patterns and temporal partitioning in a large carnivore guild. Proc. Roy Soc. B (290).

2022 Apps, P.; M. Claase; S. Lostrom; E. Van Mourik; B. Yexley; H. Webster; J. W. McNutt.  A description of a new discovery: African wild dog packs communicate with other packs by posting scent-mark messages at a shared marking site. Behavioral Ecology and Sociobiology 76(3): 37.

2022 Durant, S. Marino, A., Linnell, J.D.C., Oriol-Cotterill, A., Dloniak, S., Dolrenry, S., Funston, P., Groom, R.J., Hanssen, L., Horgan, J., Ikanda, D., Ipavec, A., Kissui, D.M., Lichtenfeld, L., McNutt, J.W., Mitchell, N., Naro, E., Samna, A., & Yirga, G. Fostering coexistence between people and large carnivores in Africa: Using a Theory of Change to identify pathways to impact and their underlying assumptions. Front. Conserv. Sci. 2:698631. doi: 10.3389/fcosc.2021.698631.

2022 Abrahms, B., Rafiq, K., Jordan, N.R., & McNutt, J.W. Long-term, climate-driven phenological shift in a tropical large carnivore. PNAS 2022 Vol. 119 No. 27 e2121667119

2022 Claase, M.J., Cherry, M.I., Apps, P.J., McNutt, J.W., Hansen, K. W. & Jordan, N.R. Inter-pack communication in African wild dogs at long-term shared marking sites. Anim.Behav. 192: 27-38

2022 Jordan, N.R., Golabek, K.A., Behr, D., Walker, R.H., Plimpton, L., Lostrom, S., Claase, M., van der Weyde, L.K., McNutt, J.W. Priority of access to food and its influence on social dynamics of an endangered Carnivore. Behav.Ecol.&Sociobiol.76(13)

2022 Jordan, N.R., Abrahms, B., Rabaiotti, D. , Rafiq, K., Woodroffe, R. & McNutt, J.W Climate change is causing endangered African wild dogs to give birth later – threatening the survival of the pack. The Conversation.

2021 Heerman, B., van Rooyen, J., Fynn, R., Biggs, D., Lewis, M., McNutt ,J.W. Husbandry and Herding: A Community-Based Approach to Addressing Illegal Wildlife Trade in Northern Botswana. Front. Conserv. Sci. 2:675493

2021  Alting, B.F.T., Golabek, K.A., Pitcher, B., McNutt, J.W., Wilson, A.M., Bates, H., & Jordan, N.R. The characteristics and consequences of African wild dog (Lycaon pictus) den site selection. Behavioral Ecology and Sociobiology 75(7). https://doi.org/10.1007/s00265-021-03047-8 

2021 Hofmann, D.; Behr, D.; McNutt, J.; Ozgul, A.; Cozzi, G. Bound within Boundaries: How well do boundaries of protected areas match movement corridors of their most mobile protected species? Journal of Applied Ecology DOI: 10.1111/1365-2664.13868

2021 Rabaiotti, D., Groom, R. McNutt, JW., Watermeyer, J., O’Neill, H.M.K., & Woodroffe, R. High temperatures and human pressures interact to influence mortality in an African carnivore. Ecology and Evolution 11(13): 8495–8506. doi: 10.1002/ece3.7601 High temperatures and human pressures interact to influence mortality in an African carnivore. Ecology and Evolution 11(13): 8495–8506. doi: 10.1002/ece3.7601

2020 Radford, C., McNutt, J.W., Rogers, T, Maslen, B. & Jordan, N. Artificial eyespots on cattle reduce predation by large carnivores. Commun Biol 3, 430.

2020 Jordan, N. Box A3: Making sense of scents: Olfactory perception in animals. Zoo Anim. Learn. & Training, 77-79.

2020 Cozzi, G. Behr, D. M., Webster, H. S., Claase, M, Bryce, C.M., Modise, B., McNutt, J.W., & Ozgul, A. African Wild Dog Dispersal and Implications for Management. J. Wildl. Management. 84,4:614-621

2020 Jordan, Neil R., Bradley P. Smith ,Robert G. Appleby, Lily M. van Eeden , & Hugh S. Webster. Addressing inequality and intolerance in human–wildlife coexistence. Conservation Biology, Volume 00, No. 0, 1–8DOI: 10.1111/cobi.13471

2020 Rafiq, K., Jordan, N.R., Meloro,C., Wilson, A., Hayward, M. & McNutt, J.W.  Scent marking strategies of a solitary carnivore: boundary and road scent marking in the leopard. Animal Behaviour 161:115-26.

2020 Behr, D. M., Cozzi, G. McNutt, J.W., & Ozgul, A. When to stay and when to leave? Proximate causes of dispersal in an endangered social carnivore. Journal of Animal Ecology 89(10): 2356–66

2020 Vitale, J. D., Jordan, N. R., Gilfillan, G. D., McNutt, J. W., & Reader, T. Spatial and seasonal patterns of communal latrine use by spotted hyenas (Crocuta crocuta) reflect a seasonal resource defense strategy. Behavioral Ecology and Sociobiology, 74(10), Article 120.

2020 Buesching, C.D. & Jordan, N.R.  Small carnivore latrines in space and time (with special emphasis on badgers and mongooses). In: Small Carnivores in Space and Time. Ed. E Do Linh San. Cambridge University Press, Cambridge UK. Part II, Chapter11.

2019 McNutt ,J.W.,  Woodroffe, R,&  Groom, R. High ambient temperatures provide an adaptive explanation for seasonal reproduction in a tropical mammal. J. Zoology 309(3): 153–60. doi:10.1111/JZO.12712

2019 Apps, P., Rafiq, K., & McNutt, J. W. Do Carnivores Have a World Wide Web of Interspecific Scent Signals?. In: Chemical Signals in Vertebrates 14 (pp. 182-202). Springer.

2019 Rafiq, K., Appleby, R.G., Edgar,J.P., Jordan, N.R., Blacker, A.R., Dexter, C.E., Jones, D.N. & Cochrane, M. OpenDropOff: an open-source, low-cost drop-off unit for animal borne devices. Methods in Ecology and Evolution 10, 1517-1522.

2019 Rafiq, K, Jordan, N.R, Wilson, A.M. , McNutt, J.W. , Hayward, M.W., Meloro, C., Wich, S.A. Golabek, K.A. Spatio-temporal factors impacting encounter occurrences between leopards and other large African predators. J. Zoology doi:10.1111/jzo.12746

2019 Rich, L., M Kelly, D Miller, D Munoz, H Robinson, JW McNutt. Sampling design and analytical advances allow for simultaneous density estimation of seven sympatric carnivore species from camera trap data. Biological Conservation: Volume 233: 12-20

2019 Kusler, A. ,N.R. Jordan , J. W. McNutt , & F. Broekhuis.  Cheetah marking trees: distribution, visitation, and behavior. African Journal of Ecology; 1–4.

2019 Chisholm,S, AB. Stein, NR. Jordan, TM. Hubel, J. Shawe-Taylor , T Fearn, JW. McNutt, A M. Wilson, S. Hailes. Parsimonious Test of Dynamic Interaction. Ecology and Evolution 9:1654–1664., doi:10.1002/ece3.4805

2019 Rafiq, K, Bryce C.M, , Rich, L.N. , Coco, C. Miller, D.A.W. , Meloro, C., Wich, S.A. , McNutt, J.W. , Hayward, M.W. DOI:10.1016/j.cub.2019.05.056 Citizen-science on safari: tourist photographs as a scalable framework for large carnivore monitoring in protected areas. Current Biology 29(14):R681

2019 Buesching, C. D., & Jordan, N. The social function of latrines: A hypothesis-driven research approach. In: Chemical Signals in Vertebrates 14 (pp. 94-103). Springer, Cham.

2018 Apps, P. , & J. W. McNutt. Are camera traps fit for purpose ? A rigorous, reproducible and realistic test of camera trap performance. African Journal of Ecology 56: 710-720.

2018 Hubel, T., Golabek, K., Rafiq, K. , McNutt, J.W., & Wilson, A.M. Movement patterns and athletic performance of leopards in the Okavango Delta. Proceedings B. Roy. Soc.B: Biological Sciences 285(1877): 20172622.

2018 Rogan, M.S., Miller, J.R.B., Lindsey, P.A., & McNutt, JW. Socioeconomic drivers of illegal bushmeat hunting in a Southern African Savanna. Biological Conservation 226: 24-31

2018 Apps, P, & J. W. McNutt. How camera traps work and how to work them. African Journal of Ecology 56:702-709.

2018 Apps, P., K. Rafiq, & J. Weldon McNutt. Camera trap records of groups of ten and eleven Honey Badgers Mellivora capensis in northern Botswana. Small Carnivore Conservation 56: i-iii.

2018 Wilson, A.M., T. Hubel, S. Wilshin, J.C. Lowe, M. Lorenc, O.P. Dewhirst, H.L.A Bartlam-Brooks, R. Diack, E. Bennitt, K.A. Golabek, R. Woledge, J.W. McNutt, N.A. Curtin, T. West. Biomechanics of predator–prey arms race in lion, zebra, cheetah and impala. Nature 554(7691): 183–88.doi:10.1038/nature25479

2017 Gilfillan G. D., McNutt, J. W., Vitale J. D.T., McComb, K. Spontaneous discrimination of urine odours in wild African lions (Panthera leo). Anim.Behav.26: 17 -185.

2017 McNutt, JW, A. B. Stein, L.P. Boggs-McNutt & N. R. Jordan. Living on the Edge: Characteristics of Human-Wildlife Conflict in a traditional livestock community in Botswana. Wildlife Research  44(7): 546–57.

2017 Walker, R.H. ,King, A.J, McNutt, JW, & N. R. Jordan. Sneeze to leave: African wild dogs (Lycaon pictus) use variable quorum thresholds facilitated by sneezes in collective decisions. Proc. Roy. Soc. B 284: 20170347

2017 Chaber, A., Cozzi, G., Broekhuis, F., Hartley, R., & McNutt , J.W. . Serosurvey for selected viral pathogens among sympatric species of the African large predator guild in northern Botswana. Journal of Wildlife Diseases 53(1):170-75

2017 Rich, L. N. Miller, D.A.W., Robinson, H.S. , McNutt, J. W.,Kelly, M.J. Carnivore distributions in Botswana are shaped by resource availability and intraguild species. J. Zoology 303(2): 90–98. doi:10.1111/jzo.12470

2017 Woodroffe, R, Groom, R, & J.W. McNutt. Hot dogs: high ambient temperatures impact reproductive success in a tropical carnivore. J. Animal Ecology 1–11.

2017 Rogan, M.S., Lindsey, P.A., Tambling, C. J., Golabek, K.A., Chase M., & McNutt, J.W. Illegal bushmeat hunters compete with predators, threatening herbivore populations in a global tourism hotspot. Biological Conservation 210: 233-242

2017 Apps, P.J, M. Claase, B. Yexley, & J.W. McNutt. Responses of wild African carnivores to the odour of 3-mercapto-3-methylbutanol, a component of wildcat and leopard urine. Journal of Ethology, 35(2), 153-159

2017 Torrents Ticó, M., Rich, L., McNutt, J.W., Nnthomiwa, M., Mothala, M., Motsamai, G. & Jordan, N.R. On the right track: comparing concurrent spoor and camera-traps surveys in Botswana. African Journal of Wildlife Research 47(2): 00–00 (October )

2017 Dewhirst, OP, K Roskilly, T Hubel, R J. Harvey, Jordan, NR; Golabek, KA, McNutt, JW.& A M. Wilson.  An exploratory clustering approach for extracting stride parameters from tracking collars on free ranging wild animals. Journal of Experimental Biology 0, 1-7 doi:10.1242/jeb.146035

2017 Jordan NR, Buse C, Wilson AM, Apps PJ, Golabek KA, Lowe JC, Van der Weyde, LK, & McNutt JW. Dynamics of direct inter-pack encounters in endangered African wild dogs. Behav. Ecol. Sociobiol. 71(8): 115. DOI : 10.1007/s00265-017-2338-9

2017 Jackson, C., R. Groom, N.R. Jordan &J.W.McNutt. The effect of relatedness and pack size on territory overlap in African wild dogs. Movement Ecol. 5:10

2017 Abrahms, B., Seidel, D.P., Dougherty, E., Hazen, E.L., Bograd, S.J., Wilson, A.M., McNutt, J.W., Costa, D.P., Blake, S., Brashares, J.S. & Getz, W.M. Suite of simple metrics reveals common movement syndromes across vertebrate taxa. Movement Ecology, 5(12): 1-11. DOI:10.1186/s40462-017-0104-2.

2016 Rich, L. N. Miller, D.A.W., Robinson, H.S., McNutt, J. W., Kelly, M.J. Extension of Cape fox distribution in Northern Botswana. Canid Biology & Conservation.19(5): 21-24. URL:

2016 Jordan, N.R., Apps, P.J., Golabek, K.A. & McNutt, J.W. Pair-specific scents in African wild dogs, Lycaon pictus, and an example of a potential method to identify signals within complex mixtures. Chap. 30, pp.461-476. In: Chemical Signals in Vertebrates 13. Ed. BA Schulte, T Goodwin & M Ferkin. New York: D. Kluwer Academic/Plenum Publishers.

2016 Hubel, T., Myatt, J., Jordan, N.J., Dewhirst, O.P., McNutt, J. W.&Alan M. Wilson. Additive opportunistic capture explains group hunting benefits in African wild dogs. Nature Communications 7:11033. DOI: 10.1038/ncomms11033.

2016 Apps, P.J. Does Deconvolution Help to Disentangle the Complexities of Mammal Odors?Chap.28, pp. 415-433. In: Chemical Signals in Vertebrates 13. Ed. BA Schulte, T Goodwin & M Ferkin. New York: D. Kluwer Academic/Plenum Publishers.

2016 Gilfillan G. D.,McNutt, J. W., Vitale J. D.T., de Iongh, H.H. & Golabek, K.A. Rare observation of the existence and masculine behaviour of maned lionesses in the Okavango Delta, Botswana. African Journal of Ecology  55(3): 383–85. doi: 10.1111/aje.12360.

2016 Abrahms, B. , N. R. Jordan, K. A. Golabek, J. W. McNutt, A. M.Wilson & J. S. Brashares. Lessons from integrating behaviour and resource selection: activity-specific responses of African wild dogs to roads. Animal Conservation1 9: 247–255.

2016 Rich, LN, Miller, DAW, Robinson, HS, McNutt, JW, and Kelly, MJ. Using Camera Trapping and Hierarchical Occupancy Modelling to Evaluate the Spatial Ecology of an African Mammal Community ed. Matt Hayward. Journal of Applied Ecology 53(4): 1225–35.doi:10.1111/1365-2664.12650.

2016 Gilfillan,G. D., Vitale, J., McNutt, J.W., D.T., McComb, K. Cross-modal individual recognition in wild African lions (Panthera leo). Biology Letters 12:20160323. URL:

2016 Abrahms, B, Sawyer, S., Jordan, N., McNutt, J.W., Wilson, A., & Brashares, J. Does wildlife resource selection accurately inform corridor conservation. Journal of Applied Ecology, 54(2): 412-422.doi: 10.1111/1365-2664.12714.

2016 Hubel, T., Myatt, J., Jordan, N.J., Dewhirst, O.P., McNutt, J. W.&Alan M. Wilson. Energy cost and return for hunting in African wild dogs and cheetahs. Nature Communications 7:11034 .DOI:10.1038/ncomms11034.

2015 Stein, A.B., Bourquin, S.L. & McNutt, J.W. Avoiding intraguild competition: Leopard feeding ecology and prey caching in Northern Botswana. African Journal of Wildlife Research 45(2):247–257

2015 Cozzi, G., Börger, L., Hutter, Abegg, D., Beran, C., McNutt, J.W, & Ozgul, A. Effect of trophy hunting leftovers on the ranging behaviour of large carnivores: A case study on spotted hyenas. PLOS One 10(3): e0121471. |DOI:10.1371/journal.pone.0121471 March 20, 2015.

2015 Apps, P. J., Weldon, P. J., & Kramer, M. Chemical signals in terrestrial vertebrates: search for design features. Natural Product Reports, 32(7), 1131-1153.

2015 Rogan, M.S., Lindsey, P.A., McNutt, J.W. Illegal bushmeat hunting in the Okavango Delta, Botswana: drivers, impacts and potential solutions. FAO Subregional Office for Southern Africa, Food and Agriculture Organization of the United Nations, Harare, Zimbabwe.68p. No.BC611

2015 Pomilia, M. A., J. W. McNutt, & N.R. Jordan. Ecological predictors of African wild dog ranging patterns in northern Botswana. Journal of Mammalogy,Volume 96, Issue 6, 24 November 2015, Pages 1214–1223

2014 Broekhuis, F., Grünewälder, S, McNutt, J.W., Macdonald, D.W. Optimal hunting conditions drive circa-lunar behaviour of a diurnal carnivore. Behavioural Ecology25 (5): 1268-1275 doi:10.1093/beheco/aru122

2014 McNutt, J W & J. Bradley. Report on Kwando (Botswana) Vulture poisoning investigation: 16 November 2013. Vulture News 66:35-41.

2014 Apps,.P.J., L. Mmualefe, N. R. Jordan, K.A. Golabek, J.W. McNutt. The “tomcat compound” 3-mercapto-3-methylbutanol occurs in the urine of free-ranging leopards but not in African lions or cheetahs. Biochemical Systematics and Ecology 53: 17–19.

2014 Jordan, N.R, Apps, P.J., Golabek, K.A.& McNutt, J.W. Top marks from top dogs: An evaluation of over marking and pair-bond advertisement in dominant pairs of African wild dogs. Animal Behaviour88: 211-217.

2013 Cozzi, G., Broekhuis, F., McNutt, J.W., Turnbull, L.A., Schmid, B. Comparison of the effects of artificial and natural barriers on large African carnivores: Implications for inter-specific relationships and connectivity.  J. Anim. Ecology, doi:10.1111/1365-2656.12039

2013 Wilson, A., J C Lowe, K Roskilly, P E Hudson, K A. Golabek, & J. W. McNutt. Locomotion dynamics of hunting in wild cheetahs. Nature, 498(13 June):185-189.

2013 Lindsey, P., et al, Illegal hunting and the bush-meat trade in African savannas: drivers, impacts and possible solutions. Biol. Conserv.160 (2013) 80–96

2013 Cozzi, G.,Broekhuis, F., McNutt, J.W. B. Schmid. Density and habitat use of lions and spotted hyenas in northern Botswana and the influence of survey and ecological variables on call-in survey estimation. Biodiversity and Conservation.(DOI) 10.1007/s10531-013-0564-7.

2013 Jordan, N.R., Golabek, K.A., Apps, P.J., Gilfillan, G.D. & McNutt, J.W. Scent marking and over marking in free-ranging packs of African wild dogs (Lycaon pictus). Ethology 119: 1– 9.

2013 McNutt, J. W. & R. Woodroffe. African wild dog. In: Mammals of Africa, Volume 5. J. Kingdon, D. Happold, T. Butynski, M. Hoffman, H. Happold & J. Kalina (Eds.). Bloomsbury, London.

2013 Packer, C. et al Conserving Large Carnivores :Dollars and Fence. Ecology Letters.Volume 16, Issue5, pages635–641, May2013

2013 Apps, P. Review: Are mammal olfactory signals hiding right under our noses? Naturwissenschaften: DOI 10.1007/s00114-013-1054-1

2013 Broekhuis, F., Cozzi, G., Valeix, M., McNutt, J.W., Macdonald, D.W. Risk avoidance in sympatric large carnivores: reactive or predictive? J. Animal Ecology May 20. doi: 10.1111/1365-2656.12077.

2012 Apps, P.& Mmualefe, L. . A versatile cryo-focussing flow switching gas chromatography inlet for trace analysis. Journal of Chromatography A, Vol.1257,(28) 2012, 171–188

2012 Prager, K.C., J.A.K. Mazet, L. Munson, S. Cleaveland, C.A. Donnelly, E.J. Dubovi, M. Szykman Gunther, R. Lines, M. Mills, H.T. Davies-Mostert, J.W. McNutt, G. Rasmussen, K.Terio, R. Woodroffe. The effect of protected areas on pathogen exposure in endangered African wild dog (Lycaon pictus) populations. Biol. Conserv.: 150:15-22.

2012 Apps, P, Mmualefe, L. & McNutt, J.W. Identification of volatiles from the secretions and excretions of African wild dogs (Lycaon pictus). J. Chemical Ecol.38(11):1450-1461

2012 Marsden, C., Woodroffe, R., Mills, M., McNutt, J., Creel, S., Groom, R., Masenga, E., Cleaveland, S., Kat, P., Rasmussen, G., Ginsberg, J., Lines, R., André, J., Begg, C., Wayne, R., & Mable, B. Spatial and temporal patterns of neutral and adaptive genetic variation in the endangered African wild dog (Lycaon pictus). Molecular Ecology: Volume 21, Issue 6, pages 1379–1393.

2012 Jackson, C., J.W. McNutt, & P. Apps. Managing the ranging behaviour of wild dogs using translocated scent marks. Wildlife Research39,31-34.

2012 Cozzi, G., Broekhuis, F., McNutt, J.W., Turnbull, L.A., Macdonald, D.W., Schmid, B. Fear of the dark or dinner by moonlight? Reduced temporal partitioning among Africa’s large carnivores. Ecology 93,(12): 2590-2599

2012 Grünewälder, S., Broekhuis,F. , D. W. Macdonald, A. Wilson, J.W. McNutt, J. Shawe-Taylor, S. Hailes. Movement activity-based classification of animal behaviour with an application to data from cheetah (Acinonyxjubatus). PLoS ONE.P 7(11): e49120 doi:10.1371/journal.pone.0049120

2012 Apps, P. , Mmualefe, L. & McNutt, J.W. A reverse engineering approach to identifying which compounds to bioassay for signalling activity in the scent marks of African wild dogs (Lycaon pictus). Proceedings of the Chemical Signals in Vertebrates XII Conference, Berlin. Aug 2011.

2012 McNutt, J. W., & M. Gusset, Declining body size in an endangered large mammal. Biol. J. Linnean Soc.:105, 8–12

2011 Webster, H., McNutt, J.W. & McComb, K. African wild dogs as a fugitive species: playback experiments investigate how wild dogs respond to their major competitors. Ethology117:1-10

2011 McNutt, J.W., Mills, M.G.L., McCreery, K., Rasmussen, G., Robbins, R. & Woodroffe, R. Lycaon pictus. In: IUCN 2011. IUCN Red List of Threatened Species.Version 2011.2.

2008 Alexander, K.A., McNutt, J.W., Briggs, M.B., Standers, P.E., Funston, P, Hemson, G., Keet, D., and van Vuuren, M. Multi-host pathogens and carnivore management in southern Africa. Comp.Immunol. Microbiol. Infect. Dis. 33, (3), Pages 249-265

2010 Alexander,K. A. & McNutt. J.W. Human behavior influences infectious disease emergence at the human–animal interface. Frontiers in Ecology and the Environment e-View. doi: 10.1890/090057

2010 Webster, H., J. W. McNutt & K. McComb. Eavesdropping and Risk Assessment Between Lions, Spotted Hyenas and African Wild Dogs. Ethology 116 233–239

2009 Gusset, M., M.J. Swarner, L. Mponwane, K. Keletile and J.W. McNutt. Human–wildlife conflict in northern Botswana: livestock predation by Endangered African wild dog Lycaon pictus and other carnivores. Oryx 43 (1):67-72. doi:10.1017/S0030605308990475

2009 Marsden, C.D., B. K. Mable, R. Woodroffe, G.S.A. Rasmussen, S. Cleaveland, M. Emmanuel, R. Thomas, J.W.McNutt, L. J. Kennedy. Highly endangered African wild dogs (Lycaon pictus) lack variation at the major histocompatibility complex. J. Heredity 100(Supplement 1): S54–65. /esp031.: S54-S65.

2009 Webster, H. Wild Dogs Vs Lions: The secret struggle for supremacy on Africa’s plains BBC Wildlife July 27 (7)

2008 McNutt, J.W. & Silk, J.B. Pup production, sex ratios, and survivorship in African wild dogs, (Lycaon pictus). Behav. Ecol. Sociobiol. 62: 1061–1067.

2008 McNutt, J.W., M. Parker, M. J. Swarner& M. Gusset. Adoption as a conservation tool for endangered African wild dogs (Lycaon pictus). South African Journal of Wildlife Research 38(2): 109–112.

2004 Boggs, L. Community-based natural resource management in the Okavango Delta. In: Rights, Resources & Rural Development : Community-based natural resource management in Southern Africa. Fabricius, C. & Koch, E.(eds.). Earthscan , London. pp147-159.

2004 Creel, S., M.G.L. Mills, J.W.McNutt. Demography and Population Dynamics of African Wild Dogs in Three Critical Ecosystems. In: The Biology & Conservation of Wild Canids, D.W. Macdonald & C. Sillero-Zubiri, eds. Oxford University Press.

2004 McNutt, J. W., M. G. L. Mills, K. McCreery, G. S. A. Rasmussen, R. Robbins, & R. Woodroffe. 2004. Lycaon pictus. Pp. 1-9 in 2004 Red List of Threatened Species (IUCN ed.). IUCN, Gland, Switzerland.

2002 McNutt, J. W. & L. P. Boggs. Another Family Dog: The social life of African wild dogs. Africa Geographic 10 (9) 38-49.

2001 McNutt, J.W. Estimating wildlife populations: Sampling and extrapolation. In: Report of Proceedings of the National Technical Predator Management and Conservation Workshop in Botswana. DWNP. October 9-12. pp54-61.

2001 McNutt, J.W. African wild dogs in northern Botswana. In: Report of Proceedings of the National Technical Predator Management and Conservation Workshop in Botswana. DWNP. October 9- 12. pp18-24.

2001 Girman, D., C. Vilà, E. Geffen, S. Creel, M. Mills, J.W.McNutt, J.R. Ginsberg, P. Kat, Mamiya & R. Wayne. Patterns of population subdivision, gene flow and genetic variability in the African wild dog (Lycaon pictus). Molecular Ecology 10 (7), 1703-1723.

2000 McNutt, J.W. Pack Power: African wild dog behaviour. BBC Wildlife 18 (2): 14 – 20.

2000 Boggs L.P. Community Power, Participation, Conflict and Development Choice: community wildlife conservation in the Okavango region of Northern Botswana. Evaluating Eden Series, Paper 17, International Institute for Environment and Development, London.

1997 Hamilton, Wm. J. III, McNutt, J.W. Determinants of conflict outcomes. Ch.6 In: Perspectives in Ethology. Vol. 12. Owings, D.H., Clark N., & Beecher, M. (eds.) Plenum Press, New York. Pp.179- 224.

1996 Osofsky, S. A., McNutt, J. W. & Hirsch, K.J. Immobilization of free-ranging African wild dogs (Lycaon pictus) using a ketamine/xylazine/atropine combination. J. Zoo Wildl. Med. 27: 528-532.

1996 McNutt, J.W. & L. P. Boggs. Running Wild: Dispelling the Myths of the African Wild Dog. Smithsonian Institution Press. Washington, D.C. 149p.

1996 McNutt, J.W. Adoption in African wild dogs (Lycaon pictus). J.Zool., Lond. 240:163-173.

1996 McNutt, J.W. Sex-biased dispersal in African wild dogs, (Lycaon pictus). Anim. Behav. 52:1067-1077.

1995 Alexander, K.A., P.W. Kat, C. House, M. K. Laurenson, J.W. McNutt, S.J. O’Brien, & B.I. Osburn. African horse sickness and African carnivores. Veterinary Microbiology 47:133-140.

1995 McNutt, J.W. Sociality and dispersal of African wild dogs (Lycaon pictus). Ph.D. Dissertation. University of California, Davis, CA

1995 Ginsberg, J.R., K.A. Alexander, S. Creel, P.W. Kat, J.W. McNutt, & M.G.L. Mills. Handling and survivorship in the African wild dog (Lycaon pictus): A survey of five ecosystems. Conserv. Biol. 9: 665-674.

1994 Alexander, K.A., N.J. MacLachlan, P.W. Kat, C. House, S.J. O’Brien, N.W. Lerche, M. Sawyer, L.G. Frank, K. Holecamp, L. Smale, J.W. McNutt, M.K. Laurenson, M.G.L. Mills, & B.I. Osburn. Evidence of natural bluetongue virus infection among African carnivores. Am J. Trop. Med. Hyg. 51: 568-576.

1993 Boggs, L.P. & J.W. McNutt. Running wild: A close look at Botswana’s painted wolves. Afr. Env. Wildl. 2: 28-36.

1992 Fuller, T. K., P.W. Kat, J.B. Bulger, A.H. Maddock, J.W. Ginsberg, R.Burrows, J.W.McNutt, & M.G.L. Mills. Population dynamics of African wild dogs. In: Wildlife 2001: Populations. D.R. McCullough & R.H. Barrett, (eds.). London. Elsevier Applied Science. Pp.1125-1138.

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